Protologue Description: Jaltomata lojae Mione, sp. nov. TYPE: ECUADOR. Prov. Loja: by guard station on the E side of Celica, 1950 m, growing along roadside, 2 may 1991, D. M. Spooner, R. Castillo, and L. López 5037 (HOLOTYPE: CONN). Figures 1 and 2.

A J. cajamarca atque J. sagasteguii Mione foliis pilis dactyliformibus, corolla sub anthesi rotata et ovarii disco lato differt.

Perennial shrub; branches and leaves densely villous, bearing finger hairs, these sometimes gland-tipped, 0.15-6 mm long, becoming less villous with age. Leaves alternate, often geminate, ovate, to 15 cm long X 6 cm wide, the margin entire to sinuate-dentate, undulate, the petiole to 4.5 cm long. Inflorescence umbelliform, to 9-flowered. Peduncle 5-9 mm long (at flowering), pedicel 8-11 mm long ( at flowering), both having a dense covering of erect finger hairs to 1.5 mm long, some hairs gland-tipped. Calyx (at flowering) light green, 9.5 mm in diameter, strongly reflexed, abaxially villous with gland-tipped finger hairs, forked hairs, and dendritic hairs all 1-2 mm long abaxially and adaxially with stalked multicellular glands 62-80 µm long (Figure 2), at fruit maturity calyx diameter 12 mm. Corolla infundibular when partially open, rotate when fully open, white with two green proximally positioned maculae straddling the radial vein to each corolla lode (Figure 1), 5 prominent lobes alternating with 5 small lobules 25-27 mm in diameter on plants we grew, 18 mm in diameter on the holotype, adaxially glabrous, abaxially with sparsely but evenly distributed stalked multicellular glands 62-80 µm (figure 2). Two classes of hairs extend out from the corolla margin: marginal hairs 110 µm long, and submarginal (attached abaxially) hairs to 0.5 mm long. Stamens 5, 5 mm long; filaments on living plants angling away from style and slightly curved outward, villous on basal ¼ of the length with unpigmented finger hairs to 1.2 mm long; anthers 2.1-2.5 mm long prior to dehiscing, 1.6-1.8 mm long after dehscing, when corolla is fully open exerted out of corolla 2 mm, otherwise included. Pollen grains 32.5 µm average diameter (n = 22 grains). Ovarian nectar disk orange (visible by eye on living flowers). Style 9-11 mm long, slender and straight ( Figure 1), exserted 3-8 mm beyond the anthers; stigma shallowly bilobed, 0.4 to 0.75 mm wide on pressed specimens, broader than the style (Figure 1), the papillae 30-60 µm long. Mature fruits orange. Seeds numerous, ovate to reniform, 1.3-1.6 mm long. Chromosome number n = 12 (nine counts).




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